Phase inverters are only present in power amps and heads with power sections.
preamps don't need them.
One of the chassis tubes is the initial gain stage after the guitar, and it's shared by all channels.
The other chassis tube is PROBABLY some kind of buffer to achieve a desired signal level.
I'm not looking at a schematic though.
I've tried a few tubes in there.
On the board, the 12At7 is supposed to be V1, and I'm loving an early 90's 12AX7 Chinese tube in there, and a new Chinese tube in the other "12AX7" position.
I tried a NOS RCA 12AT7, and it just didn't do it for me. I've also tried JJ ECC83s in both positions, and those kind of muddied it all up.
However, I'm using some JJ's in my modules, along with a few CHinese tubes.